Avella, I., E. Barajas-Ledesma, N. R. Casewell, R. A. Harrison, P.D. Rowley, E. Crittenden, W. Wüster, R. Castiglia, C. Holland, A. van der Meijden (2021) Unexpected lack of specialisation in the flow properties of spitting cobra venom. Journal of Experimental Biology (in press)


cobra on cover of Journal

Simone, Y. and A. van der Meijden (2021) Armed stem to stinger; a review of the ecological roles of the weapons of scorpions. Journal of Venomous Animals and Toxins including Tropical Diseases (in press) 

Neto, A., P. Coelho, A. van der Meijden (2021) The effect of change in mass distribution due to defensive posture on gait in fat-tailed scorpions. Journal of Comparative Physiology A 207: 117–125

DOI: 10.1007/s00359-021-01467-5

Estrada-Gómez, S., L. J. Vargas-Muñoz, A. van der Meijden, M. M. Saldarriaga-Córdoba (2021) MS/MS analysis of four scorpion venoms from Colombia, a descriptive approach. Journal of Venomous Animals and Toxins including Tropical Diseases.

DOI: 10.1590/1678-9199-JVATITD-2020-0173

Sierra Ramírez, D., G. Guevara, L. Marcela Franco Pérez, A. van der Meijden, J.C.Gonzalez Gomez, J. Carlos Valenzuela-Rojas, C. Fernando Prada Quiroga (2021) Deciphering the diet of a wandering spider (Phoneutria boliviensis; Araneae: Ctenidae) by DNA metabarcoding of gut contents. Ecology and Evolution 2021:00 (1–16)

DOI: 10.1002/ece3.7320

Valenzuela,J., J.C.Gonzalez Gomez, A. van der Meijden, J.N. Cortés, G. Guevara, L.M. Franco, S. Pekar, L.F. Garcia (2019) Prey and venom efficacy of male and female wandering spider, Phoneutria boliviensis (Araneae: Ctenidae). Toxins 11(11): 622

DOI: 10.3390/toxins11110622

González-Gómez, J.C., J.C. Valenzuela-Rojas, L.F. García, L.M. Franco Pérez3, G. Guevara, S. Buitrago, A. Cubillos, A. van der Meijden (2019) Sexual dimorphism in the biomechanical and toxicological performance in prey incapacitation of two morphologically distinct scorpion species (Chactas sp. and Centruroides sp.). Biological Journal of the Linnean Society 129(1):190–198

DOI: 10.1093/biolinnean/blz146

Grashof, D.G.B., H.M.I. Kerkkamp, S. Afonso, J. Archer, D.J. Harris, M. K. Richardson, F.J. Vonk, A. van der Meijden (2019) Transcriptome annotation and characterization of novel toxins in six scorpion species. BMC Genomics 20:645

DOI: 10.1186/s12864-019-6013-6

Wolff, J.O., Paterno, G.B, Liprandi, D., Ramirez, M.J., Bosia, F., van der Meijden, A., Michalik, P., Smith, H., Jones, B., Ravelo, A.M., Pugno, N., Herberstein, M.E. (2019) Evolution of aerial spider webs coincided with repeated structural optimization of silk anchorages. Evolution 73 (10): 2122-2134


Portik, D.M., R.C. Bell, D.C. Blackburn et al. (40 authors) (2019) Sexual dichromatism drives diversification within a major radiation of African amphibians. Systematic Biology 68(6): 859–875

DOI: 10.1101/372250

Hyperolius riggenbachi

Rasko, M. P. Coelho, Y. Simone, A. van der Meijden (2018) How to attack a scorpion: venom metering during a repeated attack. Animal Behaviour 145, 125-129.

DOI: 10.1016/j.anbehav.2018.09.019

Hadrurus arizonensis

Simone, Y., L.F. Garcia, M. Lacava, A. van der Meijden, C. Viera (2018) Predatory Versatility in Females of the Scorpion Bothriurus bonariensis (Scorpiones: Bothriuridae): Overcoming Prey with Different Defensive Mechanisms. Journal of Insect Behavior 31(4): 402–415

DOI: 10.1007/s10905-018-9677-x

Simone, Y., A. van der Meijden (2018) Fast and fine versus strong and stout: a trade-off between chela closing force and speed across nine scorpion species. Biological Journal of the Linnean Society, blx139

DOI: 10.1093/biolinnean/blx139

Chiari, Y., A. van der Meijden, A. Caccone, J. Claude & B. Gilles (2017) Self-righting potential and the evolution of shell shape in Galápagos tortoises. Scientific Reports 7: 15828

DOI: 10.1038/s41598-017-15787-7

Van der Meijden, A., B. Koch, T. van der Valk, L. J. Vargas‐Muñoz, S. Estrada‐Gómez (2017) Target‐Specificity in Scorpions; Comparing Lethality of Scorpion Venoms across Arthropods and Vertebrates. Toxins 9(10): 312.

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Wolff, J. O., A. van der Meijden, M. E. Herberstein (2017) Distinct spinning patterns gain differentiated loading tolerance of silk thread anchorages in spiders with different ecology. Proc. R. Soc. B 284: 20171124.

DOI: 10.1098/rspb.2017.1124

Coelho, P., A. Kaliontzopoulou, M. Rasko, A. van der Meijden (2017) A ‘striking’ relationship: scorpion defensive behaviour and its relation to morphology and performance. Functional Ecology.

DOI: 10.1111/1365-2435.12855

Video Abstract

scorpion strike

Estrada-Gómeza, S., L. Gomez-Ravea, L. Vargas-Muñozd, A. van der Meijden (2017) Characterizing the biological and biochemical profile of six different scorpion venoms from the Buthidae and Scorpionidae family. Toxicon.

DOI: 10.1016/j.toxicon.2017.02.007

Scorpion venom cancer cells

Van der Meijden, A., T. Kleinteich (2017) A biomechanical view on stinger diversity in scorpions. Journal of Anatomy 230(4): 497–509.

DOI: 10.1111/joa.12582

Cover Journal of Anatomy

Van der Meijden, A., P. Coelho, M. Rasko (2015) Variability in venom volume, flow rate and duration in defensive stings of five scorpion species. Toxicon 100: 60-66

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Van der Valk, T., A. van der Meijden (2014) Toxicity of scorpion venom in chick embryo and mealworm assay depending on the use of the soluble fraction versus the whole venom. Toxicon 88: 38-43


Chick embryo in eggshell

Coelho, P., P. Sousa, D.J. Harris, A. van der Meijden (2014) Deep intraspecific divergences in the medically relevant fat-tailed scorpions (Androctonus, Scorpiones). Acta Tropica 134: 43-51


Herrel, A., A. van der Meijden (2014) An analysis of the live reptile and amphibian trade in the USA compared to the global trade in endangered species. Herpetological Journal 24: 103-110

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herp trade

Pedroso, D., P. Sousa, D.J. Harris, A. van der Meijden (2013) Phylogeography of Buthus Leach, 1815 (Scorpiones: Buthidae): A multigene molecular approach reveals further complex evolutionary history in the Maghreb. African Zoology 48(2): 298–308

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Van der Meijden, A., P. Lobo Coelho, P. Sousa, A. Herrel (2013) Choose Your Weapon: Defensive Behavior Is Associated with Morphology and Performance in Scorpions. PlosONE Nov 13, 2013. DOI: 10.1371/journal.pone.0078955

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Choose your weapon

Chiari, Y., A. van der Meijden, M. Mucedda, N. Wagner, M. Veith (2013) No detection of the pathogen Batrachochytrium dendrobatis in Sardinian cave salamanders, genus Hydromantes. Amphibia-Reptilia 34: 136-141


Hydromantes ambrosii

Andreone, F., S. Loarie, A. van der Meijden, S. Nieto, M. Vences (2012) A Web Platform Implementing Information Management and Citizen-Science for the Conservation of Malagasy Amphibians. Froglog 103 (web magazine)

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Van der Meijden, A., F. Langer, R. Boistel, P. Vagovic, M. Heethoff (2012) Functional morphology and bite performance of raptorial chelicerae of camel spiders (Solifugae). Journal of Experimental Biology 215(19): 3411–3418.


Media coverage

Cover of the Journal of Experimental Biology, October 2012

Sousa, P., D.J. Harris, E. Froufe, A. van der Meijden (2012) Phylogeographic patterns of Buthus scorpions (Scorpiones: Buthidae) in the Maghreb and South-Western Europe based on CO1 mtDNA sequences. Journal of Zoology 288(1): 66-75

Map of Buthus scorpions

Van der Meijden, A., T. Kleinteich, P. Coelho (2012) Packing a pinch; Functional implications of chela shapes in scorpions using finite element analysis. Journal of Anatomy 220(5): 423-434.

Journal of Anatomy cover with Hadogenes

Chiari, Y., A. van der Meijden, M. Mucedda, J.M. Lourenco, A. Hochkirch, M. Veith (2012) Phylogeography of Sardinian cave salamanders (genus Hydromantes) is mainly determined by geomorphology. PlosOne 7(3): e32332.

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Hydromantes populations Sardinia

Kaliontzopoulou, A., D.C. Adams, A. van der Meijden, A. Perera, M.A. Carretero (2011) Relationships between head morphology, bite performance and ecology in two species of Podarcis wall lizards. Evolutionary Ecology. DOI: 10.1007/s10682-011-9538-y

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Sousa, P., E. Froufe, D.J. Harris, P. Célio Alves, A. van der Meijden (2011) Genetic diversity of Maghrebian Hottentotta (Scorpiones: Buthidae) scorpions based on CO1: new insights on the genus phylogeny and distribution. African Invertebrates. 52 (1): 135–143.

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Lötters, S., A. van der Meijden, L.A. Coloma, R. Boistel, P. Cloetens, R. Ernst, E. Lehr and M. Veith (2011) Assessing the molecular phylogeny of a near extinct group of vertebrates: the Neotropical harlequin frogs (Bufonidae; Atelopus). Systematics and Biodiversity 9(1): 45–57.

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M. Barata, A. Perera, D. J.Harris, A. van der Meijden, S. Carranza, F. Ceacero, E. García-Muñoz, D. Gonçalves, S. Henriques, F. Jorge, J. C. Marshall, L. Pedrajas, P. Sousa (2011) New observations of amphibians and reptiles in Morocco, with a special emphasis on the Eastern Region. Herpetological Bulletin 116:4-13.

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Cerastes cerastes

Van der Meijden, A., A. Crottini, J. Tarrant, A. Turner and M. Vences (2011) Multi-locus phylogeny and evolution of reproductive modes in the Pyxicephalidae, an African endemic clade of frogs. African Journal of Herpetology 2011: 1-12.

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Pyxicephalus adspersus

Lötters, S., A. van  der Meijden, D. Rödder, T. E. Koester, T. Kraus, E. La Marca, C. F.B. Haddad, and M. Veith (2010) Reinforcing and expanding the predictions of the disturbance vicariance hypothesis in Amazonian harlequin frogs: a molecular phylogenetic and climate envelope modelling approach. Biodiversity and Conservation 19: 2125-2146.

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Van der Meijden, A., A. Herrel and A. Summers (2010) Comparison of chela size and pincer force in scorpions; getting a first grip. Journal of Zoology 280: 319-325.

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Journal of Zoology cover

Chiari, Y., A. van der Meijden, O. Madsen, M. Vences, A. Meyer (2009) Base composition, selection, and phylogenetic significance of indels in the recombination activating gene-1 in vertebrates. Frontiers in Zoology, 6: 32.

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Oliehoek, P.A., P. Bijma and A. van der Meijden (2009) History and structure of the closed pedigreed population of Icelandic Sheepdogs. Genetics Selection Evolution, 41: 39

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Icelandic sheepdog

Van der Meijden, A., Y. Chiari, M. Mucedda, S. Carranza, C. Corti, and M. Veith (2009) Phylogenetic relationships of Sardinian cave salamanders, genus Hydromantes , based on mitochondrial and nuclear DNA sequence data. Molecular Phylogenetics and Evolution, 51: 399–404

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Wollenberg, K. C., D. R. Vieites, A. van der Meijden, F. Glaw, D. C. Cannatella, M. Vences (2008) Patterns of endemism and species richness in Malagasy cophyline frogs support a key role of mountainous areas for speciation. Evolution. 62 (8): 1890-1907.

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Van der Meijden, A., M. Vences, S. Hoegg, R. Boistel, A. Channing and A. Meyer. (2007) Nuclear gene phylogeny of narrow-mouthed toads (Family Microhylidae) and a discussion of competing hypotheses concerning their biogeographical origins. Molecular Phylogenetics and Evolution. 44(3): 1017–1030

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Gastrophryne carolinensis

Van der Meijden, A., R. Boistel, J. Gerlach, A. Ohler, M. Vences, A. Meyer. (2007) Molecular phylogenetic evidence for paraphyly of the genus Sooglossus, with the description of a new genus of Seychellean frogs. Biological Journal of the Linnean Society, London. 91: 347–359.

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Sooglossus skull

Van der Meijden, A. and Chiari, Y. (2006) Natrix tesselata. Marine habitat. Herpetological review 37(1):94

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Natrix tesselata

Van der Meijden, A., M. Vences, S. Hoegg and A. Meyer (2005) A previously unrecognized radiation of ranid frogs in Southern Africa revealed by nuclear and mitochondrial DNA sequences. Molecular Phylogenetics and Evolution. 37(3): 674-685.

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Pyxicephalus adspersus

Vences, M., M. Thomas, A. van der Meijden, Y. Chiari, D. R. Vieites (2005) Comparative performance of the 16S rRNA gene in DNA barcoding of amphibians. Frontiers in Zoology 2:5.

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barcode frog

Van der Meijden, A., M. Vences and A. Meyer (2004). Novel Phylogenetic relationships of the enigmatic brevicipitine and scaphiophrynine toads as revealed by sequences from the nuclear Rag-1 gene. Proceedings of the Royal Society B (Suppl.) 271: S378-S381.
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Breviceps acutirostris

Van der Meijden, A., Vredenburg, V., Sopory, A., Petirs, B., Tiwari, R., Wake, D.B.(2002) AmphibiaWeb: An information system for amphibian conservation biology. American Zoologist Vol.39 No. 5